1Mie Industry and Enterprise Support Center, Shima, Mie 517-0501, 2Graduate School of Biosphere Sciences, Hiroshima University, Higashi-Hiroshima, Hiroshima 739-8528, 3Ocean Construction Co. Ltd., Kurashiki, Okayama 711-0921, 4Fisheries Research Division, Mie Prefectural Science and Technology Promotion Center, Shima, Mie 517-0404, Japan